@article{Coulson2008,
author = {Coulson, T. and Tuljapurkar, S.},
title= {The dynamics of a quantitative trait in an age-structured population living in a variable environment},
year = {2008},
doi = {10.1086/591693},
journal = {The American Naturalist},
volume = {172},
number = {5},
pages = {599--612},
}